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Bart A. de
Cuyper
Institute for
Forestry and Game Management, 1560 Hoeilaart, Belgium
Introduction
During the last
decades, forestry in Europe has shown a growing awareness of the
high importance of a specific group of tree species known as Noble
Hardwoods. The acknowledgement of their eminent silvicultural,
economic and/or ecological values led to their upgrading from a
former status of ‘minor’ species.
Out of this
awareness arose a major concern to safeguard the existing resources
of these species (isolated individuals, relict populations,
small-scale homogeneous stands). Because one can not protect what
one does not know, substantial efforts have been needed, in the
first place, to identify the still existing resources.
Furthermore,
the gene conservation programmes, set off in different European
countries, very rapidly urged for an effective characterization of
these resources once they had been identified. In this way a varying
set of criteria, techniques and traits was developed and applied in
each country, aiming at:
determination of wild forms vis-à-vis
cultivated and hybrid forms (e.g. wild apple)
assessment of the levels of
(genetic) diversity within the entities
characterization of diversity
between geographically disjunct entities.
The EUFORGEN
Noble Hardwoods Network provides a mechanism for monitoring the
practices currently used throughout Europe for such evaluation of
genetic resources. In order to carry out an overview of the current
evaluation methods and practices in different countries, a
questionnaire was chosen as the most appropriate way to gain good
information from all the participating countries (see list of
Participants).
It was agreed
to classify the different traits used for evaluation of Noble
Hardwoods genetic resources into four categories: morphology,
phenology, resistance characteristics and biochemical/genetic
traits.
The received
information was rearranged into a fairly limited number of tables
representing the individual Noble Harwood species. The aim is to
provide each member of the Network (and other scientists concerned)
with a guide indicating where to search for additional information
in order to compare, complete, refine and/or optimize their own
evaluation techniques.
The purpose of
this task is certainly not to produce one uniform methodology for
the evaluation of Noble Hardwood species in all countries. The form
of the questionnaire used for conducting this survey allowed for
obtaining well-balanced and standard information. However, one
should keep in mind that putting the available information into a
fixed questionnaire inevitably entails a certain loss of information
which does not fit into the presented form and can not be
comprehended in the following overview tables (Tables 1-16).
Some
similarities and common features between the different countries
could be observed:
For all species considered,
the evaluation of genetic resources is most often done on the
basis of morphological and phenological traits, as these are
readily observable and do not require any sophisticated equipment
(Fig. 1). Exception is made for the Ulmus species, for
which tolerance/susceptibility to the Dutch elm disease is adopted
as a major distinctive characteristic used.
Morphological observations
most often focus on leaves and fruits/seeds (Fig. 2). Among all
phenological traits observed, flushing and flowering appear to be
the most important ones (Fig. 3).
Fig. 1. Relative importance of evaluation techniques.
Relative importance of evaluation techniques.
Fig. 2. Relative importance of morphological traits.
Relative importance of morphological traits.
Fig. 3. Relative importance of phenological traits.
Relative importance of phenological traits.
Legend for Tables 1 - 16.
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Morphology |
M1 = Leaves; M2 = Fruits and seeds; M3 =
Crown;
M4 = Stem; M5 = Bark; M6 = Growth (DBH-H);
M7 = Other |
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Phenology |
Ph1 = Flushing; Ph2 = Flowering; Ph3 =
Closure of apical bud; Ph4 = Leaf Fall; Ph5 = Autumn
colouration; Ph6 = Other |
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Biochemical/genetic traits |
BG1 = DNA analysis; BG2 = Isozymes |
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Res |
Resistance characteristics |
Table 1. Acer pseudoplatanus
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Morphology |
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Phenology |
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B/G |
Res |
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M1 |
M2 |
M3 |
M4 |
M5 |
M6 |
M7 |
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Ph1 |
Ph2 |
Ph3 |
Ph4 |
Ph5 |
Ph6 |
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BG1 |
BG2 |
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Belgium |
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Croatia |
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Czech Rep. |
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Germany |
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A |
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Netherlands |
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B |
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Romania |
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Russ. Fed. |
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Slovakia |
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A =
lamma shoots; B = buds.
Table 2. Acer campestre
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Morphology |
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Phenology |
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B/G |
Res |
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M1 |
M2 |
M3 |
M4 |
M5 |
M6 |
M7 |
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Ph1 |
Ph2 |
Ph3 |
Ph4 |
Ph5 |
Ph6 |
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BG1 |
BG2 |
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Denmark |
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Germany |
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A |
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Netherlands |
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B |
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A =
lamma shoots; B = buds.
Table 3. Acer platanoides
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Morphology† |
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Phenology‡ |
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B/G§ |
Res¶ |
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M1 |
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